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The heterotetradimers are formed subsequent the conversation of UPK2 and UPK3A moieties from the heterodimers. adhering to assembly in the endoplasmic reticulum plus the Golgi, 6 heterotetradimers organized in internal and outer rings68 variety a 16 nm AUM particle or plaque (Fig. three), packaged into DFVs and shipped to the apical membrane69.